erect, hairy herbs. Leaves opposite, petiolate or sessile, blade elliptic or
ovate, margin toothed, palmately-veined. Capitula axillary, sessile, solitary,
involucral bracts few, outer ones ciliate, inner ones globose, papillose,
enclosing ray florets, receptacle paleaceous, paleae enclosing the bisexual
florets, ligule of ray florets bidentate. Corolla of disc florets 5-lobed,
yellowish. Anther base obtuse. Styler arms longer in female florets. Cypsela compressed,
glabrous. Pappus absent.