or perennial, erect herbs. Leaves alternate, crenate, serrate or pinnatifid or
entire. Capitula heterogamous, numerous in terminal or axillary corymbs or
panicles. Bracts 2-many seriate, linear-lanceolate, receptacle flat or concave,
naked or pitted and fimbriate. Outer florets female, 2 to many seriate,
tubular, filiform or shortly ligulate, fertile, inner ones hermaphrodite,
yellow, tubular, 5-lobed. Anther base obtuse-entire. Style of female florets
much exserted, arms filiform, that of hermaphrodite florets with flattened
arms. Cypsela oblong-elliptic, pappus 1-seriate, rarely 2-seriate, hairy.