or perennial, creeping, prostrate or decumbent herbs. Leaves alternate,
pinnatifid or pinnatisect, toothed to entire. Capitula small, heterogamous,
solitary, terminal and axillary, involucre hemispherical to campanulate. Bracts
2-seriate, herbaceous, with scarious margin, receptacle conical, flat or concave.
Marginal florets female, 1-2 or many-seriate. Corolla of marginal florets
shortly conical, filiform or absent, that of hermaphrodite florets tubular,
sometimes with 2 wings, limb 4-fid. Cypsela of marginal ones stipitate,
2-ribbed or winged or wingless, pappus absent.