trees, shrubs or herbs, glabrous or densely villous. Leaves usually alternate,
petiolate or sessile, linear, 3-5 lobed, acute to acuminate, base attenuate to
auriculate, serrate to crenate, pilose to subglabrous. Capitulum solitary,
peduncled, receptacle hemispheric to convex, palea stiff, striate, usually with
2 secondary lobes enclosing the achenes, persistent. Involucral bracts 2-3
seriate, triangular. Corolla of disc-florets 5-lobed, yellow. Anthers black to
brown. Style bifid, stigmas papillate. Achene triangular to quadrangular,
black, brown or mottled, pappus of awn or of fused squamellae or absent.