Perennial or annual herbs, erect or ascending.
Stems mostly branched. Leaves opposite, rarely whorled. Flowers solitary,
terminal or axillary, or in dichasium, perfect; bracteoles present, filiform or
leaflike or absent. Calyx partly adnate to or completely free from ovary,
epigynous or hypogynous; lobes 4-6, subentire to branched. Corolla epigynous,
tubular, 4-6-merous. Stamens 4-6; filaments dilated below, glabrous or ciliate
at dilated part. Ovary inferior for corolla but from semiinferior to superior
for calyx, 3-6-locular; stigma 4-6-fid; ovules extremely numerous. Fruit a
berry. Seeds numerous, subglobose.