rarely shrubs, gigantic with mostly whorled branched. Leaves evergreen,
persistent or deciduous, stiff needle-like, arranged in 1-7 foliate fascicles.
Male and female cones on the same tree. Stamens and ovuliferous scales spirally
arranged round a common axis. Stamens many, anthers 2, naked, compact with very
short filaments. Female strobilus with scales, much enlarged, usually numerous,
accrescent. Ovules 2, collateral, bract scales also enlarged, narrow, thinner
than ovuliferous scales or reduced or hidden. Fruit a woody cone. Seeds 2,
mostly unilaterally winged.