or dioecious, evergreen shrubs or small trees, without stinging hairs. Leaves
alternate, spiral, stipules deciduous, intrapetiolar, 2-cleft; leaf blade
3-veined, margin serrulate, cystoliths punctiform. Inflorescence axillary,
cymose panicles, lateral branches usually few and short, unisexual. Glomerules
contiguous, forming dense spikes. Male flowers: perianth lobes 5, connate,
imbricate, stamens 5, filaments inflexed in bud, rudimentary ovary present.
Female flowers: perianth lobes 4 or 5, connate at the base, enlarged and fleshy
in fruits, unequal; staminodes absent. Ovary without styles, stigmas sessile,
ring-like, with villous hairs, ovules orthotropous. Achenes ovoid to obliquely
obovoid. Seeds with a little endosperm.