shrubs, monoecious or dioecious; indumentum of stellate hairs, mixed with
sessile yellow or red gland dots. Leaves alternate, subopposite, decussate,
entire or palmately lobed, palmately 3-9-nerved,
discoid almost immersed glands near base on upper surface; stipules minute.
Inflorescences terminal or axillary, simple or branched racemes or spikes. Male
flowers: 1-9 per node along rachis, sepals 3 or 4; petals and disc absent;
stamens 20-100, free, crowded on a central receptacle; disc rarely present;
pistillode minute. Female flowers: solitary or rarely paired at nodes along the
rachis; calyx spathaceous, deeply 3-6-lobed; ovary (2-) 3 (-4)-loculed,
echinate, muricate or tubercled, styles 2-3, free or shortly connate base.
Fruits capsular, (2-) 3 (-4)-loculed, dehiscing into bivalved parts, tubercled,
bristly, echinate. Seeds 3, ovoid to oblong or globose, black to red,